Wednesday, January 27, 2016

Ceratosaurus Part 1: The history of a predatory horned dinosaur.

Disclaimer: I promised you all a post about the teeth of Ceratosaurus. This is not that post. In reviewing the history of Ceratosaurus I found the post growing with background knowledge but no discussion of teeth. It make sense; there is backstory here dating back to Marsh in the 1880s! So I have decided to break this into two posts. The first post, this one, covers the history of Ceratosaurus research. The next post, out soon, will cover just the dental aspects of this charismatic animal.

Ceratosaurus, the "horned lizard," was cool beast, no doubt about it! Here we have a theropod with huge fangs that was over 20 feet long, and had bone studded armor along its back! And that is to say nothing about those flamboyant crests on its face; one sits above its nose and another one above each of its eye. It was initially described by Marsh back in 1884 and is relatively well known thanks to a number of specimens from across the American West, mainly in Colorado and east-central Utah. There are large animals, presumed to be adults, and smaller individuals (presumably juveniles) in our sample as well, so we should theoretically have a good grasp on what this animal was like. Let's take a look at the history of Ceratosaurus, starting after Marsh's initial publication.

Gilmore in 1920 redescribed Marsh's animal (thankfully!) and revised the diagnosis for Ceratosaurus. I have quoted him in full below.
Generic characters: Premaxillaries with three teeth; maxillaries with 15 teeth; dentary with 15 teeth; 9 cervical vertebrae plano-concave; dorsal vertebrae biconcave; 5 sacrals; distal caudals without special lengthening of prezygapophyses; pelvis coossified; pubis with closed obturator foramen; 4 digits in manus, first and fourth reduced; probably 3 digits in pes; dermal ossifications; abdominal ribs present.
Plate 17, Figure 1 from Gilmore (1920), showing the right side of the holotype of Ceratosaurus
Plate 17, Figure 2 from Gilmore (1920), showing the left side of the holotype of Ceratosaurus


This is a workable definition for the time but we know that some of these are not autapomorphies (things found only in one type of organism). In fact, some of these characters are ancestral for theropods, or even archosaurs. Still, we can't fault Gilmore. He was working with what comparative material he had at the time and did an excellent job. Perhaps most outstanding, even compared to papers being published in the 21st century, are his excellent illustrations of the material he is describing.

That same year (1920), Werner von Janensch published on several theropods recovered by German expeditions to the Tendaguru beds. The most famous of these is probably Elaphrosaurus bambergi, a medium sized ceratosaur subject to much speculation in the decades since. In this same publication Janensch commented on the presence of Allosaurus, which he later named Allosaurus tendagurensis (Janensch, 1925), though this has been suggested to be a carcharodontosaurid. Additionally, and relevant to the post here, he identified Ceratosaurus (?) sp., Megalosaurus (?) ingens, and Labrosaurus stechowi from the same beds.
The butt of Elaphrosaurus on display in Berlin during the 2014 SVP meeting.


Tibia, teeth, and dorsal vertebra of "Allosaurus tendagurensis", also on display in Berlin. Whatever it was, it was big.
 Janensch didn't illustrate his Ceratosaurus sp., which was based on three dorsal vertebrae, but he did illustrate his "Megalosaurus" ingens and "Labrosaurus" stechowi. These will be relevant to our discussion on the teeth of Ceratosaurus.
Figure 6 from Janensch (1920) with the tooth of  Megalosaurus (=?Ceratosaurus) ingens

Figures 7 & 8 from Janensch (1920) with a ?labial and basal view of the tooth of  Labrosaurus stechowi (=?Ceratosaurus sp.)
Work continued sporadically on the genus. In 1963 Jim Madsen and William Stokes presented at the Geological Society of America meeting in Provo about new material from the Cleveland-Lloyd Quarry in central Utah. It was obvious that there was more to this animal than had been previously described. The last (so far) phase in new Ceratosaurus species came in 2000 when Madsen and Sam Welles named two new species: C. magnicornis and C. dentisulcatus. C. magnicornis was so-named from its large nasal horn, while C. dentisulcatus derives its specific name from grooves Madsen and Welles saw on the premaxillary and anterior dentary teeth; this taxon included the remains initially described in that 1963 abstract, while C. magnicornis was named from remains found in western Colorado.

Your faithful author with part of the holotype of Ceratosaurus magnicornis at Dinosaur Journey in Fruita, Colorado
In addition to coining two new species, Madsen and Welles looked over the material that had previously been assigned to the genus by other workers, including the material from Tendaguru. This was the most comprehensive review of all the material assigned to Ceratosaurus to date. Here's a summary of their findings.
  • Ceratosaurus roechlingi (Janensch 1925) may be a very large Ceratosaurus but isn't diagnostic past Ceratosaurus sp.
  • Labrosaurus stechowi is likely a junior synonym of C. roechlingi
  • The Ceratosaurus vertebrae that Janensch (1920) identified as Ceratosaurus sp. are correctly IDed
  • Labrosaurus meriani (Janensch 1920), based on an isolated fluted tooth from the Bern Jura, in Switzerland is referred to Ceratosaurus sp.
  • Bones previously referred to Ceratosaurus from Oklahoma (Stovall, 1938) are indeterminate theropod bones at best
  • Ceratosaurus sp. teeth from western Colorado are in fact correctly IDed
  • Material collected by BYU at Dry Mesa, Colorado and Agate Basin, Wyoming, will be described soon and represent the largest known specimen of Ceratosaurus. As of 1999 the preparation of this specimen was complete.
  • Megalosaurus ingens, sometimes referred to as Ceratosaurus ingens (Rowe and Gauthier, 1990), is too big to be Ceratosaurus
  • Labrosaurus sulcatus, based on an isolated fluted tooth from the Morrison Formation of Colorado, is referred to Ceratosaurus sp.
Now some of these conclusions have held up while others have not been mentioned since 2000. Later workers, for example, don't discuss any non-North American Ceratosaurus sp. remains. There is probably a good reason for this; I will go into more detail about that on my next post. Other claims are somewhat odd (a junior synonym of a newer taxon that isn't diagnostic?). Even today, however, Madsen and Welles (2000) is the best review of all material historically assigned to Ceratosaurus.

2000 was a busy year for Ceratosaurus research. That year Brooks Britt and colleagues presented about new specimens of Ceratosaurus from Colorado, Utah, and Wyoming at the Society of Vertebrate Paleontologists annual meeting; this likely includes the material that Madsen and Welles referenced in their publication as being held at BYU. While some of this information has made its way into later publications, for the most part these specimens remain undescribed to my knowledge.

As paleontology is an evolving science, new analytical tools are always being developed. The same year that Madsen and Welles revised our view of Ceratosaurus and Britt et al. clued us in to new specimens from Wyoming, Oliver Rauhut used modern phylogenetic techniques to define Ceratosaurus as part of his Ph.D. thesis. His found the following autapomorphies (taken from Rauhut (2000) by way of Wikipedia, since I don't have access to the original thesis).
  • a narrow rounded horn core centrally placed on the fused nasals
  • a median oval groove on nasals behind horn core
  • a premaxilla with three teeth
  • premaxillary teeth with reduced extent of mesial serrations
  • chevrons that are extremely long
  • a pubis with a large, rounded notch underneath the obturator foramen
  • small epaxial osteoderms

Some of these characters still look good 16 years later, but some of them are a bit subjective, such as, "chevrons that are extremely long." This may be due to someone simplifying what Rauhut said for Wikipedia or it may reflect the long-term trend away from relative character states. Without having Rauhut's thesis I can't really say either way.

Breaking up this wall of text with a Ceratosaurus illustration. Image by DiBgd at English Wikipedia, CC BY 2.5
Rauhut (2000) wasn't the last word on the question of, "What is Ceratosaurus?" In 2008, Matt Carrano and Scott Sampson published a revised phylogentic analysis of the ceratosaurs. Since Ceratosaurus is kind of essential to understand if you're talking about a group of animals sharing its name, they came up with another revised diagnosis.
Ceratosaur with: (1) mediolaterally narrow, rounded midline horn core on the fused nasals, (2) medial oval groove on nasals behind horn core, (3) pubis with large, rounded notch underneath the obturator foramen, (4) small median dorsal osteoderms
As time passes you can see that the subjective characters have disappeared, such as the extremely long chevrons. Others characters are now better defined. We've arrived at what is essentially the definition we are using today in 2016 when we want to refer material to Ceratosaurus. Of course you may notice that this list is pretty small, meaning that most of the skeleton can't be used to identify individual bones. Carrano and Sampson (2008) do a couple of other interesting things in regards to the history of Ceratosaurus; they restrict the use of Ceratosaurus to North America (though without discussing the African material), and explain how the Madsen and Welles (2000) taxa are junior synonyms with no unambiguous autapomorphies. So from this point onward it is generally accepted that only one species of Ceratosaurus is valid, C. nasicornis, and that the genus Ceratosaurus is found only in the Morrison Formation of western North America, a point that Carrano et al. (2012) reiterate.

Juvenile Ceratosaurus partial skeleton on display at the North American Museum of Ancient Life. Photo by Zach Tirrell, CC BY-SA 2.0
Last, and most late-breaking, is a paper out today in JVP! In this paper Carrano and Choiniere discuss the arm of the holotype of Ceratosaurus from the US National Museum. The paper stays true to its subject and redescribes the arm, something the entire skeleton is in need of, as it has undergone de-mounting and additional preparation work. They found that, as many have suspected, the hand and arm of Ceratosaurus is most similar to those of early theropod dinosaurs like Dilophosaurus, and not as closely aligned with later theropods. Even from its fellow derived ceratosaurs, the Abelisauroidea, the hand and arm appear primitive - which would make sense considering its placement relative to abelisauroids in the theropod family tree. Carrano and Choiniere (2016) also show that Ceratosaurus didn't have a useless hand either. Although small and oddly shaped compared to contemporaneous Allosaurus, the hand of Ceratosaurus was still adapted to grasp items (though not to the same extent as other theropods). The lack of any preserved claws from across the Morrison associated with Ceratosaurus skeletons makes things even more difficult; was Ceratosaurus tiny-clawed, grabbing small prey items? Or did it have huge grappling-hook slashers, ready to grab on to passing sauropods? We just don't know.
Cast of the hand of the holotype of Ceratosaurus nasicornis. Note the lack of any preserved fingers. Photo by Smokeybjb, CC BY-SA 3.0
In conclusion, how can we sum up what we know of Ceratosaurus? Well there appears to be one wide-spread but relatively uncommon (compared to Allosaurus) species of Ceratosaurus that existed in western North America during the Late Jurassic. Variation that has led to different species in the Morrison Formation being named, such as Ceratosaurus magnicornis and Ceratosaurus dentisulcatus are best explained by individual variation and changes associated with the animal's growth. Other examples of Ceratosaurus may exist outside of North America but those specimens have not been rigorously examined since 2000. Numerous teeth from across western North America, Europe, and Africa have been referred to this animal, but most workers view the non-North American specimens as not part of the Ceratosaurus hypodigm. Are these referrals sound? For that, you'll have to wait until my next post!

Works Cited

Britt, Brooks, Chure, D. J., Holtz, T. R., Jr., Miles, C. A. & Stadtman, K. L. 2000. A reanalysis of the phylogenetic affinities of Ceratosaurus (Theropoda, Dinosauria) based on new specimens from Utah, Colorado, and Wyoming. Journal of Vertebrate Paleontology 20: 32A

Carrano, Matthew T., Roger BJ Benson, and Scott D. Sampson. 2012. "The phylogeny of Tetanurae (Dinosauria: Theropoda)." Journal of Systematic Palaeontology 10.2: 211-300

Carrano, Matthew T.  & Jonah Choiniere 2016. New information on the forearm and manus of Ceratosaurus nasicornis Marsh, 1884 (Dinosauria, Theropoda), with implications for theropod forelimb evolution. Journal of Vertebrate Paleontology

Carrano, Matthew T., and Scott D. Sampson. 2008. "The phylogeny of Ceratosauria (Dinosauria: Theropoda)." Journal of Systematic Palaeontology 6.02: 183-236.

Gilmore, Charles W. 1920. Osteology of the carnivorous Dinosauria in the United States National Museum, with special reference to the genera Antrodemus (Allosaurus) and Ceratosaurus. Bulletin of the United States National Museum 110: 1–154.

Janensch, Werner. "Über Elaphrosaurus bambergi und die megalosaurier aus den Tendaguru-Schichten Deutsch-Ostafrikas." Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin 8 (1920): 226-235.

Janensch, Werner. 1925. "Die Coelurosaurier und Theropoden der Tendaguru-Schichten Deutsch-Ostafrikas". Palaeontographica, Supplement 7 1: 1–99.

Madsen, Jim H. Jr., and Stokes, William L., 1963, New information on the Jurassic dinosaur Ceratosaurus: Geological Society of America, Special Paper 73, p. 90 (abs.)


Madsen, Jim H.; Welles, Samuel P. 2000. Ceratosaurus (Dinosauria, Theropoda): A Revised Osteology. Utah Geological Survey. pp. 1–80.

Marsh, O.C. 1884. "Principal characters of American Jurassic dinosaurs, part VIII: The order Theropoda" American Journal of Science 27(160): 329–340
Rauhut, Oliver. 2000. The interrelationships and evolution of basal theropods (Dinosauria, Saurischia). Ph.D. dissertation, Univ. Bristol [U.K.]. 440 pp

Rauhut, Oliver W. M. 2011. "Theropod dinosaurs from the Late Jurassic of Tendaguru (Tanzania)". Special Papers in Palaeontology 86: 195–239.

Rowe, T., and Jacques Gauthier. 1990. "Ceratosauria." in The Dinosauria, Weishampel, Dodson, and Osmólska, eds. University of California Press. pp. 151-168.
Stovall, J. Willis. 1938. "The Morrison of Oklahoma and its dinosaurs." The Journal of Geology: 583-600.

Wednesday, January 20, 2016

"Why does science have to name every little thing?"

One of the most common questions I got when I was teaching as some variation on that theme. "Why do scientists have to make things so complicated?" "Why can't they just call it something simple?" It is a question bigger than just high school level as well; one of the barriers to effective science communication and education seems to be the general "fear" of overly technical language . The general public seems to view scientists as speaking in convoluted and complex terms.

It is true that scientists have devised some very complex ways of describing things that might seem simple to a layman. And sometimes we can get wrapped up in using the terms we are familiar with when talking about our research, to the detriment of any non-technical audience. The media is also partially to blame as well, with perpetuating ideas such as all prehistoric reptiles are dinosaurs. But why do these terms exist in the first place?

This brings me to Hendrickx et al. (2015). Hendrickx and his coauthors published a paper last year breaking down theropod dinosaur teeth and analyzing many different aspects of their morphology. In addition, the authors created a standardized terminology for future paleontologists to use when describing their dinosaur teeth. Hendrickx and his coauthors explain why creating such a terminology is needed succinctly; in effect answering the question of why scientists create names for so many things.
...several pivotal theropod taxa with well-preserved dentitions still lack a thorough dental description...leading numerous authors to identify isolated theropod teeth to broad clades with uncertainty...isolated teeth are key pieces of evidence to assess vertebrate paleoecological diversity and are often used for stable isotopic studies with various applications...A better understanding of theropod anatomy and morphological variation is therefore central to help resolving systematic relationships and to provide paleoecological clues. Tooth morphology is tied to diet, which has extensive evolutionary repercussions, such as morphological convergence, more than other parts of the skeleton. Yet, theropod teeth have been shown to possess many diagnostic features of taxonomic value...Although theropod teeth seem simple at first sight, this is effectively a result of the absence of comprehensive studies on tooth anatomy and morphological variation among theropods, as well as the lack of a uniform anatomical nomenclature.
What does the wall of text mean? Basically, theropod dinosaur teeth can be used to study evolutionary relationships, paleoecology, and several other important things in paleontology, but no one has bothered to come up with a good way to talk about them.

That's the crux of scientific terminology; coming up with a good way to talk about things. Good, in this case, means usable. Terms should describe well-defined parts of an organism's anatomy. If we say, "the tip of the tooth," on a tooth that has multiple "tips", how are we to know which specific tip we're talking about? Are there differences between "wrinkles" and "grooves?"

Figure 1 from Hendrickx et al., 2015

Without understanding the distinctions between subtle anatomical differences in different taxa, how are we to find out if these features are actually taxonomically important? A quick example from the Morrison Formation. Here in western Colorado we have basically two relatively common large theropods from the Morrison: Allosaurus and Ceratosaurus. Skeletal remains of Allosaurus, including teeth, seem to dominate in the Morrison making up 3/4 of all the theropod remains (Foster, 2007). Teeth attributed to Ceratosaurus do turn up in the field, however, and are usually distinguished by the presence of ridges near their bases. Now that we have Hendrickx et al.'s paper, we can go into a bit more depth. We can say, for instance, that the teeth attributed to Ceratosaurus have basal fluting, and these flutes are not seen in the contemporaneous Allosaurus. So this may help us distinguish between these teeth in the field and keeps us from mistaking Ceratosaurus teeth (with their flutes) with wrinkled or ornamented teeth (or tooth fragments).

The description of Ceratosaurus teeth by previous authors, however, has been lacking in detail and confusing, often using different terms for the same anatomical feature. As Hendrickx et al. note, having their framework in place will help facilitate such a description and they specifically mention Ceratosaurus as being in need of such a redescription. Hopefully such a project will be forthcoming. This topic will also be the focus of my next blog post!

Going forward I am hoping to see a theropod-wide tooth catalog. While Hendrickx et al. do point out that teeth are quick to change, evolutionarily speaking, to changes in diet and feeding behavior, they also note the taxonomic utility of teeth. While many theropod teeth can't be narrowed down to a genus or species, being able to address higher-level taxonomic questions with teeth is important. In addition, some taxa appear to have diagnostic dental modifications. Doing systematic studies and descriptions of theropod teeth may yield more information on what characters are taxonomically useful and potentially add autapomorphies to established genera.

My biggest complaint is that the authors did not examine what a theropod tooth is. They identify problems with past work, the utility of teeth, and the need for a framework but there is no way to determine if this framework is applicable to a given tooth. Obviously for teeth attached to theropod jaws this isn't a problem, but the majority of the dental fossil record for archosaurs consists of isolated shed teeth. While workers in the Cretaceous and Jurassic strata have this problem to a lesser degree (though it is possible that some crocodylomorphs developed similar tooth morphologies), those of us working in the Triassic are confronted with a host of dental convergences! One need look no further than the saga of Revueltosaurus to find examples of teeth that look similar between widely divergent clades. In the Triassic there are plenty of carnivorous reptiles, many with laterally compressed teeth. While in truth the terms developed by Hendrickx et al. (2015) are likely to be broadly applicable, a brief discussion of what synapomorphies exist among the dentition of theropods would have been appreciated, so that those of us working under all that overburden could sort our rauisuchian teeth from our dinosaur teeth just a little easier.

Works Cited

Foster, John. 2007. "Allosaurus fragilis". Jurassic West: The Dinosaurs of the Morrison Formation and Their World. Bloomington, Indiana: Indiana University Press. pp. 170–176

Hendrickx, C., Mateus, O. and Araújo, R., 2015. A proposed terminology of theropod teeth (Dinosauria, Saurischia). Journal of Vertebrate Paleontology,35(5), p.e982797. 


Wednesday, January 13, 2016

Ch-ch-changes!

So things have been slow at the pub here for a bit because several changes have occurred since our last post. I will summarize them here quickly!

1.  Gary Vecchiarelli, one of the founders of the Prehistoric Pub, has decided to step down from blogging, at least temporarily. That leaves just me (Rob) onboard as the sole blogger for the Prehistoric Pub. Consequently, posts may come slower than in the past. Be assured, however, that the Pub is not and will not be abandoned. Upcoming topics will be the theropod tooth paper review that was promised to you (and I'm sure you are all anxiously awaiting), as well as coverage of the Utah Friends of Paleontology annual meeting, coming up at the start of April in Moab - more details as that gets closer.

2. At the same time as Gary was departing the Pub, I as undergoing major life changes. I changed jobs and changed states and am now working at museum in Colorado. I love the area and job but the move has left little time for blogging. Now that I am settling in, expect blog posts to resume. I am shooting for two a month (not including this introductory one!) for 2016. One will probably be technical in nature while the other may be a reflection on general trends or topics in paleontology. And since this is the Prehistoric Pub, I will probably throw in a beer post or two throughout the year.

Cheers, and happy 2016 everyone!

Friday, October 23, 2015

When is a Triassic fauna not Triassic?

In recent years among paleontologists who work on the Triassic/Jurassic boundary there has been some serious excitement about a new locality in northeastern Utah that hosts a wide variety of cool fossils. It has been named the Saint's and Sinners Quarry and has been actively worked by crews from Brigham Young University in Provo, Utah since 2009. Based on abstracts and news articles it is clear that the fauna is diverse and well represented by multiple specimens. Having been at SVP in recent years I have been able to see images of the fossils coming out of the quarry first hand. Over 11,500 fossils have been removed from the quarry which Brooks Britt (from BYU) and others estimate is only 33% excavated. Virtually all of the fossils are preserved in 3D, allowing us to have spectacular insights into animals we do not have much data from, due to crushing and other concerns. Most of the specimens are even articulated! My hat is off to all of the BYU and Dinosaur National Monument crews who have been literally working on the edge of a cliff to extract these remains.

But. You knew there was a "but" coming, didn't you? But while the fossils themselves are spectacular there has been a trend in the last couple years to refer to this bone bed as being Late Triassic in age.  Admittedly aeolian deposits are hard to date; they tend to lack any significant ash deposits and detritial zircons (which can be used to constrain ages in other sedimentary rocks) are not really useful in sand dunes. That is what the Nugget Sandstone is - a deposit of windblown sand in western North America that began during the latest Triassic Period and persisted well into the Early Jurassic (see Sprinkel et al., 2011 for more details). This sand sea expanded as paleolatitude changed and western North America drifted further away from the equator and into the "dry belt" where warm, arid climatic conditions exist. This pattern can even be seen in the Late Triassic Chinle Formation at Dinosaur National Monument, as presented on at SVP this year (Irmis et al., 2015).

The first reports of the quarry (Chambers et al., 2011) suggested that Britt and colleagues at first assigned an Early Jurassic age to the deposit. This date was keeping with the general consensus that the Triassic/Jurassic boundary was somewhere within the Nugget. By 2012, however, it appeared that the teams views changed. That year Engelmann and others (note -the actual abstract doesn't appear to be available any longer) presented an abstract at the GSA conference in Charlotte, NC. In the title they state that a new drepanosaur has been found in the Nugget Sandstone and state that it has biostratigraphic importance. They also explicitly question the Jurassic age of the Nugget (they literally put a question mark in front of the word Jurassic) based on this new find. This new drepanosaur is pretty dang cool! The team expanded on it in recent SVP meetings (Chure et al., 2013; Chure et al., 2015). This critter seems to show highly derived characters shared only with Drepanosaurus (a European form) that indicate it was a specialized fossorial (digging) animal. The kicker here is that all other known drepanosaurs come from definitive Triassic strata. The Nugget drepanosaur comes from a quarry 55 meters above the last reliably dated strata (the Bell Canyon Formation, which sits between the Chinle and Nugget in northeastern Utah).

So what's the problem? Well this year the team again presented on some more spectacular fossils from the Saints and Sinners Quarry, including a large toothed pterosaur that is very closely related to the Early Jurassic European pterosaur Dimorphodon (Britt et al., 2015). This story has been picked up by the national media who have been reporting this site as being Late Triassic in age. Let's do a quick review of the evidence for a Late Triassic age.

Evidence of a Triassic Age of the Saints and Sinners Quarry

  • Presence of a drepanosaur
  • Presence of several small sphenosuchians
  • In a formation that is traditionally considered to span the Triassic/Jurassic Boundary

Okay...that's not really a convincing list. This is especially true if you are claiming that this extraordinary interdunal wetland deposit represents a Triassic assemblage unlike any other in western North America. In fact two of the "pros" can actually be taken as a "con" and the third I think is ambiguous.
Allow me to present a list of why I have concerns about a Triassic age for this quarry.

Why the Saints and Sinners Quarry may be Jurassic in age

  • In a formation that is traditionally considered to span the Triassic/Jurassic Boundary
  • Quarry located 55 meters above the last Triassic-dated rocks (~1/2 the thickness of the Nugget)
  • Presence of the most-derived drepanosaur yet discovered
  • Presence of a pterosaur that is most similar to a Jurassic pterosaur
  • Presence of a medium-large bodied theropod in the quarry in addition to a coelophysoid
  • Presence of several small sphenosuchians
  • No phytosaurs
  • No aetosaurs
  • No metoposaurs
  • Upper Nugget lacks a Triassic ichnofauna
Well, does this mean case closed? No. While my list may be longer it isn't the final word on anything. Several of these points rely on the absence of taxa like phytosaurs and we all know that the absence of evidence is not the evidence of absence. Still, taken as a suite of things, I am not convinced that this quarry is Triassic. There are a few ways that perhaps we could do to see if I'm wrong.

  • Phylogenetic analysis of the sphenosuchians - closely related to Chinle or Kayenta taxa?
  • Phylogenetic analysis of new drepanosaur compared to the still-unnamed Ghost Ranch form
  • Phylogenetic analysis of the theropods - are they closer to Coelophysis or later taxa?
  • Additional fieldwork to look for unambiguous biostratigraphic markers
To me this fauna looks like a typical Early Jurassic fauna from western North America with a drepanosaur thrown in. Could it be an impoverished Late Triassic fauna that also has several highly derived taxa in it? I suppose and I will happily eat my hat if that is the case. What a great collection of Triassic taxa it would be! With the data that have been presented thus far I just can't see it though.

Why does this matter? Timing is everything in evolution. One of the big ways we as paleontologists talk about paleobiogeography is in terms of dispersal and vicariance. Are animals (and plants, and fungi, etc.) slowly moving into new areas or are populations split up by new barriers, isolating groups that then adapt in their own directions? To put it in the context of the Nugget fossils, are we seeing evidence that many disparate clades were widespread in the Late Triassic, or are we seeing similar taxa from elsewhere in North America in the Early Jurassic adapting to new environments? These questions have serious implications for our understanding of the rate of evolution among all these groups. By tying down the date of the Saints and Sinners Quarry we will be better able to answer some of these questions.

Final caveat: this is all based off of abstracts, talks, and posters and conferences, some of which I was unable to attend or access (this is why people should archive their conference presentations on FigShare - but I digress). I am extremely excited to see the peer reviewed publications that should result from these finds. And it may very well be that their method for dating the quarry is more nuanced than they have already presented. As always, I suppose, "Wait for the paper."



Works Cited
Britt, B. B., Chure, D., Engelmann, G., Dalla Vecchia, F., Scheetz, R. D., Meek, S., Thelin, C., Chambers, M. A NEW, LARGE, NON-PTERODACTYLOID PTEROSAUR FROM A LATE TRIASSIC INTERDUNAL DESERT ENVIRONMENT WITHIN THE EOLIAN NUGGET SANDSTONE OF NORTHEASTERN UTAH, USA INDICATES EARLY PTEROSAURS WERE ECOLOGICALLY DIVERSE AND GEOGRAPHICALLY WIDESPREAD. Journal of Vertebrate Paleontology, Program and Abstracts, 2015 p. 97

Chure, D. J., Andrus, A. S., Britt, B. B., Engelmann, G. F., Pritchard, A. C., Scheetz, R., Chambers, M. MICRO CT IMAGERY REVEALS A UNIQUE MANUS MORPHOLOGY WITH DIGGING/SCRATCHING ADAPTATIONS IN THE SAINTS AND SINNERS QUARRY (SSQ) DREPANOSAUR, NUGGET SANDSTONE (LATE TRIASSIC), NORTHEASTERN UT Journal of Vertebrate Paleontology, Program and Abstracts, 2015 p. 107

Chure, D., Britt, B., Engelmann, G., Andrus, A., Scheetz, R. DREPANOSAURS IN THE DESERT: MULTIPLE SKELETONS OF A NEW DREPANOSAURID FROM THE EOLIAN NUGGET SANDSTONE (?LATE TRIASSIC - EARLY JURASSIC), SAINTS AND SINNERS QUARRY, UTAH: MORPHOLOGY, RELATIONSHIPS, AND BIOSTRATIGRAPHIC IMPLICATIONS Journal of Vertebrate Paleontology, Program and Abstracts, 2013 p. 106

Chambers, Mariah, Hales Kimberly, Brooks B. Britt, Daniel J. Chure, George F. Engelmann, and Rod Scheetz. "Preliminary taphonomic analysis of a Ceolophysoid theropod dinosaur bonebed in the Early Jurassic Nugget Sandstone of Utah." In Geological Society of America Abstracts with Programs, vol. 42, no. 4, p. 16. 2011.

Engelmann, G., Britt, B., Chure, D., Andrus, A., Scheetz, R. MICROVERTEBRATES FROM THE SAINTS AND SINNERS QUARRY (NUGGET SANDSTONE: ?LATE TRIASSIC–EARLY JURASSIC): A REMARKABLE WINDOW ONTO THE DIVERSITY AND PALEOECOLOGY OF SMALL VERTEBRATES IN AN ANCIENT EOLIAN ENVIRONMENT  Journal of Vertebrate Paleontology, Program and Abstracts, 2013 p. 122

Engelmann, George F., Daniel J. Chure, Brooks B. Britt, and Austin Andrus. "The biostratigraphic and paleoecological significance of a new drepanosaur from the Triassic-? Jurassic Nugget Sandstone of northeastern Utah." In 2012 GSA Annual Meeting in Charlotte. 2012.

Irmis, R. B., Chure, D. J., Wiersma, J. P. LATITUDINAL GRADIENTS IN LATE TRIASSIC NONMARINE ECOSYSTEMS: NEW INSIGHTS FROM THE UPPER CHINLE FORMATION OF
NORTHEASTERN UTAH, USA Journal of Vertebrate Paleontology, Program and Abstracts, 2015 p. 149

Sprinkel, Douglas A., Bart J. Kowallis, and Paul H. Jensen. "Correlation and age of the Nugget Sandstone and Glen Canyon Group, Utah." Utah Geological Association Publication 40 (2011): 131-149.

Sunday, September 13, 2015

Dr. Thomas Holtz is my George Clooney.

When people ask me what I would do if I met this or that celebrity, I always say the same.  "I don't get starstruck." I really don't.  Being a bartender for so many years in popular spots, I have met my fair share of celebrities.  However, believe it or not, I get more giddy meeting amazing people in the field of science.  These celebrities are rockstars in my book.  Not only are they cool, but they inspire future generations with awesomeness.  Dr. Thomas Holtz is my George Clooney.  Why George Clooney Gary?  Well, just like George Clooney screams Hollywood, Dr. Holtz screams science and paleontology. The world needs more celebrities in the field of education.  Dr. Holtz is one of those celebrities.

Hats off to you Dr. Holtz and Happy Birthday.  Thank you for being an inspiration to us all.  I haven't posted in a great while, so I thought what better way to clean off the rust than to honor a great paleontologist.  I took the summer off to work hard, venture off on two field classes, and try to relax before going into another year of classes.  Special thank you to Lisa Buckley and Robert Gay for contributing to this site.  You are good friends and this site is yours also.  The pub is about sharing science and promoting good friends, so anything I can do, I do my best to help others.

P.S.  My son is a big fan of Dr. Holtz and not long ago made a character of him in Lego's video game Jurassic World along with others.  Click here to see!  Thought I'd share it again if you haven't seen it.  It is truly an epic creation.